Contrasts of Vegetation.—Placed immediately upon the Equator and surrounded by extensive oceans, it is not surprising that the various islands of the Archipelago should be almost always clothed with a forest vegetation from the level of the sea to the summits of the loftiest mountains. This is the general rule. Sumatra, New Guinea, Borneo, the Philippines and the Moluccas, and the uncultivated parts of Java and Celebes, are all forest countries, except a few small and unimportant tracts, due perhaps, in some cases, to ancient cultivation or accidental fires. To this, however, there is one important exception in the island of Timor and all the smaller islands around it, in which there is absolutely no forest such as exists in the other islands, and this character extends in a lesser degree to Flores, Sumbawa, Lombock, and Bali.

In Timor the most common trees are Eucalypti of several species, also characteristic of Australia, with sandalwood, acacia, and other sorts in less abundance. These are scattered over the country more or less thickly, but, never so as to deserve the name of a forest. Coarse and scanty grasses grow beneath them on the more barren hills, and a luxuriant herbage in the moister localities. In the islands between Timor and Java there is often a more thickly wooded country abounding in thorny and prickly trees. These seldom reach any great height, and during the force of the dry season they almost completely lose their leaves, allowing the ground beneath them to be parched up, and contrasting strongly with the damp, gloomy, ever-verdant forests of the other islands. This peculiar character, which extends in a less degree to the southern peninsula of Celebes and the east end of Java, is most probably owing to the proximity of Australia. The south-east monsoon, which lasts for about two-thirds of the year (from March to November), blowing over the northern parts of that country, produces a degree of heat and dryness which assimilates the vegetation and physical aspect of the adjacent islands to its own. A little further eastward in Timor and the Ke Islands, a moister climate prevails; the southeast winds blowing from the Pacific through Torres Straits and over the damp forests of New Guinea, and as a consequence, every rocky islet is clothed with verdure to its very summit. Further west again, as the same dry winds blow over a wider and wider extent of ocean, they have time to absorb fresh moisture, and we accordingly find the island of Java possessing a less and less arid climate, until in the extreme west near Batavia, rain occurs more or less all the year round, and the mountains are everywhere clothed with forests of unexampled luxuriance.

Contrasts in Depth of Sea.—It was first pointed out by Mr. George Windsor Earl, in a paper read before the Royal Geographical Society in 1845, and subsequently in a pamphlet "On the Physical Geography of South-Eastern Asia and Australia", dated 1855, that a shallow sea connected the great islands of Sumatra, Java, and Borneo with the Asiatic continent, with which their natural productions generally agreed; while a similar shallow sea connected New Guinea and some of the adjacent islands to Australia, all being characterised by the presence of marsupials.

We have here a clue to the most radical contrast in the Archipelago, and by following it out in detail I have arrived at the conclusion that we can draw a line among the islands, which shall so divide them that one-half shall truly belong to Asia, while the other shall no less certainly be allied to Australia. I term these respectively the Indo-Malayan and the Austro-Malayan divisions of the Archipelago.

On referring to pages 12, 13, and 36 of Mr. Earl's pamphlet, it will be seen that he maintains the former connection of Asia and Australia as an important part of his view; whereas, I dwell mainly on their long continued separation. Notwithstanding this and other important differences between us, to him undoubtedly belongs the merit of first indicating the division of the Archipelago into an Australian and an Asiatic region, which it has been my good fortune to establish by more detailed observations.

Contrasts in Natural Productions.—To understand the importance of this class of facts, and its bearing upon the former distribution of land and sea, it is necessary to consider the results arrived at by geologists and naturalists in other parts of the world.

It is now generally admitted that the present distribution of living things on the surface of the earth is mainly the result of the last series of changes that it has undergone. Geology teaches us that the surface of the land, and the distribution of land and water, is everywhere slowly changing. It further teaches us that the forms of life which inhabit that surface have, during every period of which we possess any record, been also slowly changing.

It is not now necessary to say anything about how either of those changes took place; as to that, opinions may differ; but as to the fact that the changes themselves have occurred, from the earliest geological ages down to the present day, and are still going on, there is no difference of opinion. Every successive stratum of sedimentary rock, sand, or gravel, is a proof that changes of level have taken place; and the different species of animals and plants, whose remains are found in these deposits, prove that corresponding changes did occur in the organic world.

Taking, therefore, these two series of changes for granted, most of the present peculiarities and anomalies in the distribution of species may be directly traced to them. In our own islands, with a very few trifling exceptions, every quadruped, bird, reptile, insect, and plant, is found also on the adjacent continent. In the small islands of Sardinia and Corsica, there are some quadrupeds and insects, and many plants, quite peculiar. In Ceylon, more closely connected to India than Britain is to Europe, many animals and plants are different from those found in India, and peculiar to the island. In the Galapagos Islands, almost every indigenous living thing is peculiar to them, though closely resembling other kinds found in the nearest parts of the American continent.

Most naturalists now admit that these facts can only be explained by the greater or less lapse of time since the islands were upraised from beneath the ocean, or were separated from the nearest land; and this will be generally (though not always) indicated by the depth of the intervening sea. The enormous thickness of many marine deposits through wide areas shows that subsidence has often continued (with intermitting periods of repose) during epochs of immense duration. The depth of sea produced by such subsidence will therefore generally be a measure of time; and in like manner, the change which organic forms have undergone is a measure of time. When we make proper allowance for the continued introduction of new animals and plants from surrounding countries by those natural means of dispersal which have been so well explained by Sir Charles Lyell and Mr. Darwin, it is remarkable how closely these two measures correspond. Britain is separated from the continent by a very shallow sea, and only in a very few cases have our animals or plants begun to show a difference from the corresponding continental species. Corsica and Sardinia, divided from Italy by a much deeper sea, present a much greater difference in their organic forms. Cuba, separated from Yucatan by a wider and deeper strait, differs more markedly, so that most of its productions are of distinct and peculiar species; while Madagascar, divided from Africa by a deep channel three hundred miles wide, possesses so many peculiar features as to indicate separation at a very remote antiquity, or even to render it doubtful whether the two countries have ever been absolutely united.

Returning now to the Malay Archipelago, we find that all the wide expanse of sea which divides Java, Sumatra, and Borneo from each other, and from Malacca and Siam, is so shallow that ships can anchor in any part of it, since it rarely exceeds forty fathoms in depth; and if we go as far as the line of a hundred fathoms, we shall include the Philippine Islands and Bali, east of Java. If, therefore, these islands have been separated from each other and the continent by subsidence of the intervening tracts of land, we should conclude that the separation has been comparatively recent, since the depth to which the land has subsided is so small. It is also to be remarked that the great chain of active volcanoes in Sumatra and Java furnishes us with a sufficient cause for such subsidence, since the enormous masses of matter they have thrown out would take away the foundations of the surrounding district; and this may be the true explanation of the often-noticed fact that volcanoes and volcanic chains are always near the sea. The subsidence they produce around them will, in time, make a sea, if one does not already exist.

But, it is when we examine the zoology of these countries that we find what we most require—evidence of a very striking character that these great islands must have once formed a part of the continent, and could only have been separated at a very recent geological epoch. The elephant and tapir of Sumatra and Borneo, the rhinoceros of Sumatra and the allied species of Java, the wild cattle of Borneo and the kind long supposed to be peculiar to Java, are now all known to inhabit some part or other of Southern Asia. None of these large animals could possibly have passed over the arms of the sea which now separate these countries, and their presence plainly indicates that a land communication must have existed since the origin of the species. Among the smaller mammals, a considerable portion are common to each island and the continent; but the vast physical changes that must have occurred during the breaking up and subsidence of such extensive regions have led to the extinction of some in one or more of the islands, and in some cases there seems also to have been time for a change of species to have taken place. Birds and insects illustrate the same view, for every family and almost every genus of these groups found in any of the islands occurs also on the Asiatic continent, and in a great number of cases the species are exactly identical. Birds offer us one of the best means of determining the law of distribution; for though at first sight it would appear that the watery boundaries which keep out the land quadrupeds could be easily passed over by birds, yet practically it is not so; for if we leave out the aquatic tribes which are pre-eminently wanderers, it is found that the others (and especially the Passeres, or true perching-birds, which form the vast majority) are generally as strictly limited by straits and arms of the sea as are quadrupeds themselves. As an instance, among the islands of which I am now speaking, it is a remarkable fact that Java possesses numerous birds which never pass over to Sumatra, though they are separated by a strait only fifteen miles wide, and with islands in mid-channel. Java, in fact, possesses more birds and insects peculiar to itself than either Sumatra or Borneo, and this would indicate that it was earliest separated from the continent; next in organic individuality is Borneo, while Sumatra is so nearly identical in all its animal forms with the peninsula of Malacca, that we may safely conclude it to have been the most recently dismembered island.

The general result therefore, at which we arrive, is that the great islands of Java, Sumatra, and Borneo resemble in their natural productions the adjacent parts of the continent, almost as much as such widely-separated districts could be expected to do even if they still formed a part of Asia; and this close resemblance, joined with the fact of the wide extent of sea which separates them being so uniformly and remarkably shallow, and lastly, the existence of the extensive range of volcanoes in Sumatra and Java, which have poured out vast quantities of subterranean matter and have built up extensive plateaux and lofty mountain ranges, thus furnishing a vera causa for a parallel line of subsidence—all lead irresistibly to the conclusion that at a very recent geological epoch, the continent of Asia extended far beyond its present limits in a south-easterly direction, including the islands of Java, Sumatra, and Borneo, and probably reaching as far as the present 100-fathom line of soundings.

The Philippine Islands agree in many respects with Asia and the other islands, but present some anomalies, which seem to indicate that they were separated at an earlier period, and have since been subject to many revolutions in their physical geography.